Lullula arborea
TAXONOMY
Alauda arborea Linnaeus, 1758, Europe-Sweden.
OTHER COMMON NAMES
French: Alouette lulu; German: Heidelerche; Spanish: Totovнa.
PHYSICAL CHARACTERISTICS
5.9 in (15 cm); male 0.7–1.2 oz (21–35 g); female 1.1–1.2 oz
(30–35 g); smaller and more slender than skylark and crested
lark. Plumage buff brown, upperparts and chest streaked, distinguished
from other larks by broad, white supercilium which
continues to nape. Black-and-white pattern of alula (first digit
of wing) feathers very conspicuous. Crown feathers can be
raised to a small crest.
DISTRIBUTION
Northern West Africa, Europe from the Mediterranean to
southern Scandinavia, Asia Minor east to Iran and Turkmeniya.
HABITAT
Requires habitat with short grass for feeding, higher vegetation
for nesting, exposed trees or bushes as song-perches.
BEHAVIOR
Not gregarious, even during migration. Northern populations
migratory, wintering in southern Europe and North Africa;
southern populations mostly resident. Arboreal, walks on
branches and perches on treetops, bushes, and wires. For
song-flight, males take off from perch in tree top, ascend at an
angle, spiral upward and fly circles on nearly same level
165–330 ft (50–100 m) above ground, singing all the time (unpaired
males fly higher than paired ones). Descent either in
stages while still singing or sometimes silently with wings
closed. Song-flight takes two minutes on average, but unpaired
males have been watched singing for 70 and 94 minutes.
Males also sing from perch or ground, and frequently on
moonlit nights. Song composed of pleasing and soft phrases,
hesitant at the beginning, then increasingly forceful and
louder. High site-fidelity of males has been shown by ringing
experiments.
FEEDING ECOLOGY AND DIET
Feeds on small insects and spiders during breeding season, otherwise
mainly granivorous.
REPRODUCTIVE BIOLOGY
Monogamous. Breeds March through June; cup-shaped nest
built by female. Two, sometimes three broods annually; clutch
size normally three to five eggs, female incubates for 11–15
days. Young fed by both parents, leave nest after eight days before
being able to fly. If female starts incubation of second
brood, male cares for young of first clutch alone.
CONSERVATION STATUS
Not threatened, though declining populations in Europe
caused by habitat loss; listed in Annex I of the European Birds
Directive.
SIGNIFICANCE TO HUMANS
None known.
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