There are two kinds of crabs. The irregular-tailed
crabs, or Anomurans, include the hermit, porcelain,
and mole crabs. Some have a small, tapered
abdomen, and others, especially the hermit crabs,
have a somewhat coiled abdomen that is tucked
into a discarded snail shell. The true crabs, or
Brachyurans (short-tailed crabs), have a greatly
reduced abdomen that is tucked forward beneath
the carapace.
Contrary to popular belief, crabs do not always
scuttle sideways; they can walk forward or backward
with equal facility. Afew of the fast moving
crabs seem to use two pairs of legs when running.
The blue crab (Callinectes sapidus) family, which
also includes the familiar lady crab (Ovalipes
ocellatus) have a paddlelike last pair of appendages
and are fast and agile swimmers.
Crabs vary greatly in size and shape. The
smallest are pea crabs, which measure only a few
millimeters across the carapace, while the legs of a
Japanese spider crab can span up to four meters.
The differences in size and shape have allowed
crabs to colonize a wide range of habitats. Most of
the 4,500 species of true crabs are marine, inhabiting
the rocky intertidal zone or burrowing in sand
and mud. Some species are benthic and others invade
freshwater, especially in the tropics. Still others
are terrestrial or semiterrestrial, often ranging
far away from water, although they generally prefer
moist places. A tree-dwelling tropical species
feeds on coconuts.
Ecologically, almost all crabs function as scavengers,
feeding on dead or dying animals or organic
debris of any kind. Many species are also active
predators, capable of killing small fish and
breaking open the shells of different mollusks.
Porcelain crabs, for example, have one of their
mouthparts equipped with long, feathery setae
that are swept through the water. They catch small
particles and transfer them to other parts of the
mouth, where they are eaten. Organic matter is
dismembered by the pincers and conveyed to the
mouth by a series of appendages called maxillipeds
and maxillae.Apair of comblike mandibles
guard the mouth and chop the food into tiny
pieces, which are then swallowed.Ashort esophagus
leads to a gastric mill analogous to the gizzard
of other animals, where food is ground up.
Crab Reproduction
Reproductive behaviors vary, but during courtship
males may use various chemical, visual, or
acoustic signals including pheromones, waving
chelipeds back and forth, or drumming at the entrance
to the female's burrow. Other males build
pyramid-shaped mounds to attract females. Following
courtship the male transfers sperm to her
in a packet called a spermatophore. The eggs are
attached to the female's abdominal appendages
and form an egg mass between the abdomen and
the thorax. The abdomen is therefore out of place
and hangs downward when the female is carrying
eggs. The eggs are usually brightly colored with
carotenoid pigments, and can be red, yellow, blue,
brown, or green. They range in size and number
between species, but a large marine crab can lay
up to a million eggs at a time. The tiny larvae are
called zoea and are very common summer components
of marine plankton. It takes one or two
more molts before the crablike form is evident.
Many of the semiterrestrial and freshwater species
develop directly into miniature crabs.
Crabs as Food
Crabs are food for a variety of other animals, including
fish and larger invertebrates. To protect
themselves, some species carefully place sea
anemones on their carapace, while others, such as
the decorator crab, glue pieces of shell all over
their carapace. Still other species, such as the pea
crab, take refuge inside oysters and other shellfish,
while boring crabs dig burrows into mud
substrates or soft limestone rocks.
Classification:
Kingdom: Animalia
Phylum: Arthropoda
Subphylum: Crustacea
Class: Malacostraca
Order: Decapoda
Families: Homaridae or Nephropsidae (true lobsters),
Palinuridae (spiny lobsters or sea crayfish),
Scyllaridae (slipper, Spanish, or shovel
lobsters), Polychelidae (deep-sea lobsters);
Brachyryncha (true crabs), Anomura (irregular-
tailed crabs)
Geographical location: All the earth's oceans, except
at the poles
Habitat: Mostly marine, although some crab species
live much of their lives on land near salt
water
Special anatomy: Segmented exoskeleton; large
claws on front pair of legs (one often larger
than the other); eyes on the ends of stalks
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